Diversification dynamics of darkling beetles. A, Pattern of diversification rates through time of tenebrionids. The lineage-through-time plot is reconstructed with 1,000 random post-burn-in trees (the LTT based on the median values is also figured in red). The results of diversification analyses (net diversification, speciation and extinction rates) are portrayed and indicated the shift during the Cretaceous terrestrial revolution (KTR). B, Overview of the possible correlates of diversification for tenebrionids on a global scale. A synthetic view is provided for: (i) the floristic turnovers during the KTR -; (ii) climate cycles over times; (iii) climatic conditions over time; (iv) changes in global average temperatures (data compiled from various sources -); and (v) biome evolution over time (paleomaps redrawn from Blakey ). A typical arid-adapted tenebrionid (Onymacris unguicularis) is also included for illustration purpose.