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Table 1 Family frequencies, fitnesses, and change in frequency of competitive allele, full-sib families, no policing. From left to right: Family types, frequencies, frequency of offspring genotypes, allele frequency within families, mean offspring competitive intensity, family fitness, and change in frequency of competitive allele due to selection within the family.

From: The evolution of competition and policing: opposing selection within and among groups

Family Freq Offspring Genotypes Family Mean
   AA Aa Aa qi. zi. = ziFS wi. Δqi.
AA × AA p4 1 --- --- 0 z0 = z1FS 1-z1FS 0
AA × Aa 4p3q 1/2 1/2 --- 1/4 z0+za/2 = z2FS 1-z2FS za/8z2FS
AA × aa 2p2q2 --- 1 --- 1/2 z0+za = z3FS 1-z3FS 0
Aa × Aa 4p2q2 1/4 1/2 1/4 1/2 z0+za = z4FS 1-z4FS za/4z4FS
Aa × aa 4pq3 --- 1/2 1/2 3/4 z0+3za/2 = z5FS 1-z5FS za/8z5FS
aa × aa q4 --- --- 1 1 z0+2za = z6FS 1-z6FS 0