Phylogenetic tree inferred from maximum likelihood (ML) analysis based on ITS sequences. Only LB and PB over 70%, and PP over 0.90 were reported. Thickened branches indicate LB/PB between 90%–100% and PP between 0.95–1.0. Thickened branches with dots on the root nodes represent 100% PB/LB and 1.0 PP values. PSR species were identified according to the GCPSR criterion, the Dist. species were recognised by the variations of ITS sequences, and the Finally accepted species were defined by integrating molecular phylogenetic analyses with morphological studies. Species previously allocated in the section Phalloideae and later nested in the section Lepidella are highlighted by asterisks after their names. Sequences from type collections are indicated by (T).