Patch depletion, niche structuring and the evolution of co-operative foraging

Table 2 Evolvable parameters of individuals

Category	Parameter	St. dev.	Min	Max	Units
Grouping
	n_R(tolerated neighbors)	0.2(15)	0.0	-	individuals
Zones	z _R	0.2(15)	0	-	m
	z _A	0.2(15)	z _R	-	m
Angles	a _R	0.2(360)	0.0	360	degrees
	a _A	0.2(360)	0.0	360	degrees
Foraging
Durations	t_M(MOVE)	0.2(2)	1	18	t _MIN
	t_F(FOODSCAN)	0.2(2)	1	18	t _MIN
Distances	d_M(MOVE)	0.2(15)	0.0	-	m
	d_F(FOODSCAN)	0.2(15)	0.0	-	m
Angles	a_M(MOVE)	0.2(360)	0.0	360	degrees
	a_F(FOODSCAN)	0.2(360)	0.0	360	degrees
Probabilities	p_M(MOVE after MOVE)	0.2	0.0	-
	p_SE(FOODSCAN after EAT)	0.2	0.0	-
	p_SN(FOODSCAN after NOFOOD)	0.2	0.0	-
	p_MTF(MOVETOFOOD)	0.2	0.0	-

The value of mutated genes is drawn from a normal distribution about the mother's gene value. The standard deviation of this normal distribution is scaled (0.2(x)) relative to what is considered a reasonable range for the parameter. The maximum of durations is imposed due to how the model was programmed, but is high enough not to affect the results. t_MINis the minimal time interval (see Section 1.1. in Additional file 1 ).

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