Fig. 4

Difference in gamma statistic values between reconstructed and simulated trees, where simulated trees were generated under three models with different relationships between rates of molecular evolution and speciation rates. Negative scores indicate that reconstructed trees have nodes distributed more towards the root (‘tippier’) than the simulated tree. The three simulation models are Unlinked (instantaneous covariance of molecular rates and speciation rates = 0), Continuous (instantaneous covariance = 0.0044), and Punctuated (instantaneous covariance = 0, bursts of substitutions added at speciation events). Topologies and node times were reconstructed using three different methods, an uncorrelated lognormal (UCLN) rate prior in BEAST 2, an autocorrelated lognormal rate (ACLN) prior in PAML, and the UCLN in PAML