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Table 2 New tribal concepts and their genera

From: Multigene phylogenetic analysis redefines dung beetles relationships and classification (Coleoptera: Scarabaeidae: Scarabaeinae)

 

Tribe/Genera

Inv.

Biogeographic Region

Tribe Sisyphini sensu novo

  

1

Epirinus Reiche, 1841

M, P

Afrotropical

2

Neosisyphus Muller, 1942

M, P

Afrotropical, Oriental

3

Nesosisyphus Vinson, 1946

L

Mauritius

4

Sisyphus Latreille, 1807

P

Afrotropical, Oriental, Palaearctic, Neotropical

Tribe Dichotomiini sensu novo

  

1

Chalcocopris Burmeister, 1846

M

Neotropical

2

Dichotomius Hope, 1838

M, P

Neotropical, Nearctic

3

Holocephalus Hope, 1838

S

Neotropical

4

Isocopris Pereira & Martinez, 1960

S

Neotropical

Tribe Coprini sensu novo

  

1

Copris Muller, 1764

M, P

Afrotropical, Oriental, Palaearctic, Nearctic, Neotropical

2

Litocopris Waterhouse, 1891

P

Afrotropical

3

Microcopris Balthasar 1958

M

Oriental

4

Pseudocopris Ferreira, 1960

L

Afrotropical

5

Pseudopedaria Felsche, 1904

M

Afrotropical

Tribe Deltochilini (Canthonini) sensu novo

  

1

Anisocanthon Martinez & Perreira, 1956

S

Neotropical

2

Anomiopus Westwood, 1842

M, P

Neotropical

3

Canthon Hoffmansegg, 1817

M, P

Neotropical, Nearctic

4

Canthonidia Paulian, 1939

S

Neotropical

5

Canthotrypes Paulian, 1939

S

Neotropical

6

Deltepilissus Pereira, 1949

S

Neotropical

7

Deltochilum Eschscholtz, 1822

M, P

Neotropical, Nearctic

8

Eudinopus Burmeister, 1840

P

Neotropical

9

Hansreia Halffter & Martinez, 1977

M, P

Neotropical

10

Holocanthon Martinez & Pereira, 1956

S

Neotropical

11

Malagoniella Martinez, 1961

M, P

Neotropical

12

Megathopa Eschscholtz, 1822

P

Neotropical

13

Megathoposoma Balthasar, 1939

M, P

Neotropical

14

Melanocanthon Halffter, 1958

S

Nearctic

15

Pseudocanthon Bates, 1887

S

Neotropical, Nearctic

16

Scatonomus Erichson, 1835

M

Neotropical

17

Scybalocanthon Martinez, 1948

M, P

Neotropical

18

Scybalophagus Martinez, 1953

M, P

Neotropical

19

Sylvicanthon Halffter & Marttinez, 1977

M

Neotropical

20

Tetraechma Blanchard, 1843

M

Neotropical

21

Vulcanocanthon Pereira & Martinez, 1960

S

Neotropical

22

Xenocanthon Martinez, 1952

S

Neotropical

  1. List of genera assigned to the redefined tribes based on their new concepts. Column "Inv." (investigation source) specifies evidence based on which genus was attributed to the tribe. Abbreviations are as follows: (M) morphological phylogeny [1], (P) present phylogeny, (S) synapomorphies or diagnostic characters checked (material examined per genus is given in Additional file 24: Table S6), (L) synapomorphies or diagnostic characters were not investigated and genus was attributed based on description and overall similarity to the type genus of tribe