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Table 1 The state of segment specific characters in mid-body segment and the pygidium in P. dumerilii

From: A metameric origin for the annelid pygidium?

Character

Body segment

Pygidium

Outer annulus

Present

Present

Appendages

One pair of biramous parapodia with each two cirri and two chaetae bundles

One pair of cirri

Ventral ganglia

Paired ganglia interconnected by a commissure

Absent, but a pygidial commissure is present

Peripheral ganglia

One pair of parapodial ganglia

One pair of presumably cirral ganglia

Muscular elements

Longitudinal muscle bundles, transverse muscles, parapodial muscle complexes

Circular muscles, specific ventral muscle complex

Blood vascular system

Dorsal and ventral longitudinal vessels, lateral and parapodial vessels, gut plexus

Circular blood sinus

Secondary cavity

Paired coelomic sacs delineated by myoepithelial cells formed by schizocoely

Unpaired coelomic cavity delineated by myoepithelial cells, of unclear developmental origin

Free swimming cells in the secondary cavity

Numerous different coelomocytes

Absent

Nephridia

One pair of metanephridia

Absent