Collection sites and distribution areas of Orcula species. The three maps show the distributions of all Orcula species. Black dots mark the location of sample sites investigated during this study. (A) Distribution ranges O. dolium, O. jetschini, O. wagneri and O. schmidtii, and O. zilchi. The dotted line marks the area enlarged in Figures B and C. Map modified from . (B and C) Distribution ranges Orcula species in the Alps.