Maximum-likelihood tree and nested clade TCS networks based on combined dataset. Nodal values above the line indicate bootstrap supports and poster probabilities of MP/ML/BI, while the values under the line are the divergence times estimated by MDIV. Colours represent geographical groups and spatterworks stand for locations of sample sites. The black dots refer to missing steps intermediate between observed haplotypes. Nested clades are indicated by 'N-#', where N is the nesting level and # is the number of individuals assigned to the clades within each level.