N

Population size

ϕ

Phenotype of an individual

ϕ
^{o}

Mean phenotype of an individual's offspring

δ

ϕ^{o} ϕ

$\widehat{\overline{\delta}}$

Expected mean value of δ in the population

w

Fitness of an individual

$\tilde{w}$

Expected fitness in the current environment

f
_{
ϕ
}

Frequency of phenotype ϕ in the population

Ω

$\frac{w}{\overline{w}}$ conditional on $\overline{w}$ ≠ 0

H($\overline{w}$)

Harmonic mean of $\overline{w}$

μ_{
i
}(w)

i^{th}central moment of w

$\overline{X}$ or Ave (X)

Average value of X in population

$\widehat{X}$ or E(x)

Expected value of random variable X
