| |
4-NP-α-Glc
|
VP
|
ADH
|
ODC
|
SAC
|
RAF
|
CEL
|
ARA
|
CIT
|
MR
|
ADO
|
SOR
|
LDC
|
H2S
|
|---|
|
Cronobacter spp.
|
+
|
+
|
+
|
+
|
+
|
+
|
+
|
+
|
+
|
-
|
-
|
-
|
-
|
-
|
|
Buttiauxella agrestis
|
v
|
-
|
-
|
+
|
-
|
+
|
+
|
+
|
+
|
+
|
-
|
-
|
-
|
-
|
|
Citrobacter koseri
|
-
|
-
|
v
|
+
|
v
|
-
|
+
|
+
|
+
|
+
|
+
|
+
|
-
|
-
|
|
Citrobacter freundii
|
-
|
-
|
v
|
-
|
v
|
v
|
v
|
+
|
v
|
+
|
-
|
+
|
-
|
+
|
|
Edwardsiella tarda
|
-
|
-
|
-
|
+
|
-
|
-
|
-
|
-
|
-
|
+
|
-
|
-
|
+
|
+
|
|
Enterobacter aerogenes
|
-
|
+
|
-
|
+
|
+
|
+
|
+
|
+
|
+
|
-
|
+
|
+
|
+
|
-
|
|
Enterobacter asburiae
|
-
|
-
|
v
|
+
|
+
|
v
|
+
|
+
|
+
|
+
|
-
|
+
|
-
|
-
|
|
Enterobacter cancerogenus
|
-
|
+
|
+
|
+
|
-
|
-
|
+
|
+
|
+
|
-
|
-
|
-
|
-
|
-
|
|
Enterobacter cloacae
|
-
|
+
|
+
|
+
|
+
|
+
|
+
|
+
|
+
|
-
|
v
|
+
|
-
|
-
|
|
Enterobacter gergoviae
|
-
|
+
|
-
|
+
|
+
|
+
|
+
|
+
|
+
|
-
|
-
|
-
|
+
|
-
|
|
Enterobacter hormaechei
|
-
|
+
|
v
|
+
|
+
|
-
|
+
|
+
|
+
|
v
|
-
|
-
|
-
|
-
|
|
Enterobacter pyrinus
|
v
|
v
|
-
|
+
|
+
|
-
|
+
|
+
|
-
|
v
|
-
|
-
|
+
|
-
|
|
Enterobacter helveticus*
|
+
|
-
|
-
|
-
|
-
|
-
|
+
|
+
|
-
|
+
|
-
|
-
|
-
|
-
|
|
Enterobacter turicensis*
|
+
|
-
|
-
|
-
|
-
|
-
|
+
|
+
|
-
|
+
|
-
|
-
|
-
|
-
|
|
Escherichia coli
|
-
|
-
|
v
|
v
|
v
|
v
|
-
|
+
|
-
|
+
|
-
|
+
|
(+)
|
-
|
|
Hafnia alvei
|
(-)
|
(+)
|
-
|
+
|
-
|
-
|
(-)
|
+
|
-
|
v
|
-
|
-
|
+
|
-
|
|
Klebsiella pneumoniae
|
(-)
|
+
|
-
|
-
|
+
|
+
|
+
|
+
|
+
|
-
|
+
|
+
|
+
|
-
|
|
Kluyvera spp.
|
v
|
-
|
-
|
+
|
+
|
+
|
+
|
+
|
(+)
|
+
|
-
|
v
|
v
|
-
|
|
Leclercia adecarboxylata
|
-
|
-
|
-
|
-
|
v
|
v
|
+
|
+
|
-
|
+
|
+
|
-
|
-
|
-
|
|
Morganella morganii
|
-
|
-
|
-
|
+
|
-
|
-
|
-
|
-
|
-
|
+
|
-
|
-
|
-
|
(-)
|
|
Pantoea spp.
|
-
|
v
|
-
|
-
|
v
|
v
|
v
|
+
|
v
|
v
|
-
|
v
|
-
|
-
|
|
Proteus vulgaris
|
+
|
-
|
-
|
v
|
(+)
|
-
|
-
|
-
|
v
|
v
|
-
|
-
|
-
|
+
|
|
Providencia spp.
|
-
|
-
|
-
|
-
|
v
|
-
|
-
|
-
|
v
|
+
|
v
|
-
|
-
|
v
|
|
Rahnella aquatilis
|
-
|
-
|
v
|
-
|
+
|
+
|
+
|
+
|
(-)
|
-
|
-
|
+
|
-
|
-
|
|
Raoultella terrigena
|
-
|
+
|
-
|
(-)
|
+
|
+
|
+
|
+
|
v
|
v
|
+
|
+
|
+
|
-
|
|
Salmonella sv.
|
-
|
-
|
v
|
(+)
|
-
|
-
|
v
|
(+)
|
v
|
+
|
-
|
v
|
(+)
|
v
|
|
Serratia marcescens
|
v
|
+
|
-
|
+
|
+
|
-
|
-
|
-
|
+
|
(-)
|
v
|
+
|
+
|
-
|
|
Yersinia enterocolitica
|
-
|
-
|
-
|
+
|
+
|
-
|
v
|
+
|
-
|
+
|
-
|
+
|
-
|
-
|
- 4-NP-α-Glc, metabolism of 4-NP-α-glucoside; VP, Voges-Proskauer; ADH, arginine dihydrolase; ODC, ornithine decarboxylase; SAC, acid from sucrose; RAF, acid from raffinose; CEL, acid from cellobiose; ARA, acid from arabinose; CIT, use of citrate as sole carbon source (Simmon's); ADO, acid from adonitol; SOR, acid from sorbitol; LDC, lysine decarboxylase; MR, methly red test; H2S, production of hydrogen sulphide. +, 90–100% positive; (+), 80–90% positive; v, 20–80% positive; (-), 10–20% positive; -, less than 10% positive. Data was derived from this study and from Manual of Clinical Microbiology, 7th Edition [25]. *Enterobacter helveticus and E. turicensis are novel Enterobacter species [38].
