Maps of selected paths in AGGREG illustrating the main findings. The major conclusion is that butterflies made tortuous paths (small moves interrupted by frequent and large turns) when flying within habitat (path 1), but flew straighter (long moves, small turns) in the matrix (paths 4, 5 and 6). Path 2 shows the tendency to return into the patch after having crossed the habitat patch boundary . Path 3 resulted from a spontaneous flight (without experimental manipulation) and illustrates that experimental artefacts due to manipulation were very unlikely the cause for straighter paths in the matrix (see Methods for details). Path 4 illustrates the return to a suitable habitat patch, with the last turn at a distance of c. 25 m from the patch, well around the estimate of perceptual range. Grey: habitat patches; +: release site in the matrix.