The numbers represent the benefit/cost ratio above which selfishness is favored (expression 5), and the cost/benefit ratio below which altruism is favored (expression 6). Low values are more selfish. Each column defines one kind of colony structure and each row represents selection on a different kind of gene, the first three in a female offspring and the last in a queen. The "Maternal" conditions are for when a mother would be selected to imprint one of her daughters to perform the action in question. Fathers would be selected to imprint daughters under the patrigenic conditions.