Figure 5

Diagnostic for hitch-hiking. Circles and bars indicate median and interquartile ranges from all combinations of recurrent sweeps with τ=10³,10⁴, s _b =10⁻³,10⁻², u=10⁻⁵,10⁻⁶and β=0.25,0.5,1,2. Parameter combinations which give a median ${â}_B<0$ are denoted with an open circle whereas simulation conditions which result in ${â}_B>0$ are shown with closed circles. The x-axis is the relative excess of non-synonymous substitutions due to linked positive selection, calculated as $({\bar{k}}^{\prime }-\bar{k})/\bar{k}$ averaged across all time points after 40000 generations, where $\bar{k}$ is the mean number of non-synonymous substitutions at non-beneficial sites averaged across 500 simulations with no positive selection and the prime indicates the corresponding values in a simulation with both positive and negative selection.