From: The role of sexual preferences in intrasexual female competition
 |  | Males | |||||||||||||||
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Females | Â | x1 | x2 | x3 | x4 | x5 | x6 | x7 | x8 | x9 | x10 | x11 | x12 | x13 | x14 | x15 | x16 |
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 | X4 |
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 | x8 |
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 | x9 |
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 | x10 |
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 | x11 |
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 | x13 | (x13x1) | (x13x2 | (x13x3) | (x13x4) | (x13x5) | (x13x6) | (x13x7) | (x13x8) | (x13x9) | (x13x10) | (x13x11) | (x13x12) | (x13x13) | (x13x14) | (x13x15) | (x13x16) |
 | x14 | (x14x1) | (x14x2) | (x14x3) | (x14x4) | (x14x5) | (x14x6) | (x14x7) | (x14x8) | (x14x9) | (x14x10) | (x14x11) | (x14x12) | (x14x13) | (x14x14) | (x14x15) | (x14x16) |
 | x15 | (x15x1) | (x15x2) | (x15x3) | (x15x4) | (x15x5) | (x15x6) | (x15x7) | (x15x8) | (x15x9) | (x15x10) | (x15x11) | (x15x12) | (x15x13) | (x15x14) | (x15x15) | (x15x16) |
 | x16 | (x16x1) | (x16x2) | (x16x3) | (x16x4) | (x16x5) | (x16x6) | (x16x7) | (x16x8) | (x16x9) | (x16x10) | (x16x11) | (x16x12) | (x16x13) | (x16x14) | (x16x15) | (x16x16) |
 | Z1 = αp(x1 + x5 + x9 + x13) + α(x2 + x3 + x6 + x7 + x10 + x11 + x14 + x15) + x4 + x8 + x12 + x16 | ||||||||||||||||
 | Z2 = α(x1 + x2 + x5 + x6 + x9 + x10 + x13 + x14) + x3 + x4 + x7 + x8 + x11 + x12 + x15 + x16 | ||||||||||||||||
 | Z3 = α(x1 + x3 + x5 + x7 + x9 + x11 + x13 + x15) + x2 + x4 + x6 + x8 + x10 + x12 + x14 + x16 |