Characters | Pronotocupedini | Notocupedini | Lithocupedini | Brochocoleini | Ommatini | Tetraphalerini | |
---|---|---|---|---|---|---|---|
1 | Pentagonal enlarged window punctures along elytral suture | present | absent | absent | absent | absent | absent |
2 | Ratio of length of 3rd and 4th segments of antenna | equal in length | equal in length | 3rd longer than 4th antennomere | 3rd longer than 4th antennomere | 3rd longer than 4th antennomere | 3rd longer than 4th antennomere |
3 | Size of window punctures enlarged, separated by very distinct rib-like ridges | present | present | absent | absent | absent | absent |
4 | Abdominal ventrites | overlapping on fore border of next | overlapping on fore border of next | overlapping on fore border of next | flat | flat | flat |
5 | Gular sutures | complete | complete | complete | incomplete | incomplete | absent |
6 | Epipleural rim | one row of cells | one row of cells | one row of cells | more than one row of cells | one row of cells | one row of cells |
7 | Median epicranial suture on the dorsal head surface | present | present | present | present | absent | present |
8 | The ventral head surface with lateral antennal grooves | absent | absent | absent | absent | absent | present |