New fossil species of ommatids (Coleoptera: Archostemata) from the Middle Mesozoic of China illuminating the phylogeny of Ommatidae

Table 2 Evolutionary trends of key characters for Ommatidae

	Characters	Pronotocupedini	Notocupedini	Lithocupedini	Brochocoleini	Ommatini	Tetraphalerini
1	Pentagonal enlarged window punctures along elytral suture	present	absent	absent	absent	absent	absent
2	Ratio of length of 3^rd and 4^th segments of antenna	equal in length	equal in length	3^rd longer than 4^th antennomere	3^rd longer than 4^th antennomere	3^rd longer than 4^th antennomere	3^rd longer than 4^th antennomere
3	Size of window punctures enlarged, separated by very distinct rib-like ridges	present	present	absent	absent	absent	absent
4	Abdominal ventrites	overlapping on fore border of next	overlapping on fore border of next	overlapping on fore border of next	flat	flat	flat
5	Gular sutures	complete	complete	complete	incomplete	incomplete	absent
6	Epipleural rim	one row of cells	one row of cells	one row of cells	more than one row of cells	one row of cells	one row of cells
7	Median epicranial suture on the dorsal head surface	present	present	present	present	absent	present
8	The ventral head surface with lateral antennal grooves	absent	absent	absent	absent	absent	present

ISSN: 2730-7182