New phylogeny of the Grumichellini, Leptorussini and Triplectidinae, presented with morphological characteristics. Phylogeny based on the composite BI tree in Figure 5. The morphological characters tibial spur formula, forewing, hind wing, male and female genitalia, respectively, are presented next to the individual species or genera. Species names highlighted in bold are those contrasting with previous classifications (see text for detail). The triangle shows the placements of Symphitoneuria species and the star that of Triplectidina species. Colours of taxon names follow Figure 3.