Unrooted consensus network consistent with optimal Bayesian and MP topologies shown in Fig. 2. Branch lengths are proportional to number of changes obtained in MP tree #2 [see Additional file 1] derived from the combined dataset, using assigned branch length in PAUP and assuming accelerated transformations (ACCTRAN). Dashed lines for Amblysomus are diagrammatic only. Squares indicate the two root positions (i.e., where Tenrecidae attaches) favored by 4 of 8 optimal MP trees (Fig. 2B), stars those favored by the majority of Bayesian post-burnin topologies (Fig. 2A). The branch leading to Tenrecidae is 106 steps; the longest intra-chrysochlorid branches are those leading to Chrysospalax and Eremitalpa at 15 steps each. Circles denote nodes to which the 100 randomly generated outgroup taxa were attracted, and are proportionately larger when more random outgroups attach at that node (indicated by the number within/adjacent to each circle). Seven of the randomly generated outgroups did not yield a resolved position for the root.